What's the dif between red and high rufus chocolate agouti?

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Thank you so much for posting this. I am fascinated by this. I would love to read the research, is it available anywhere?
I attached the link to the paper in my post but in case you missed it, here it is again:
A composite six bp in-frame deletion in the melanocortin 1 receptor (MC1R) gene is associated with the Japanese brindling coat colour in rabbits (Oryctolagus cuniculus)

This is the bit - highlighted - where they say they can't find a difference between ED and ES, except there is one other allele there (aside from e and ej):
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